Education

PhD Biochemistry, University of Cambridge, Cambridge, United Kingdom, 1980.
M. Phil. Biochemistry, University of Cambridge, Cambridge, United Kingdom, 1978.
MS Biology, American University of Beirut, Beirut, Lebanon, 1977.
BS Biology, University of Jordan, Amman, Jordan, 1971. (with distinction)

Professional Experience

1991- Professor, Department of Biology, American University of Beirut, Beirut, Lebanon.
2001-2002 Sabbatical leave, Institute of Biomedical & Life Sciences, University of Glasgow, United Kingdom.
1987-1993 Chairman, Department of Biology, American University of Beirut, Beirut, Lebanon.
1986-1991 Associate Professor, Department of Biology, American University of Beirut, Beirut, Lebanon.
1986-1988 Part-time Lecturer, Department of Natural Sciences, The Lebanese University, Beirut, Lebanon.
1985-1986 Associate Professor, Department of Biological Sciences, Yarmouk University, Irbid, Jordan.
1985-1986 Director of Personnel, Yarmouk University, Irbid, Jordan.
1983-1986 Assistant to the President, Yarmouk University, Irbid, Jordan.
1984-1985 Director, Department of Planning and Development, Yarmouk University, Irbid, Jordan.
1980-1985 Assistant Professor, Department of Biological Sciences, Yarmouk University, Irbid, Jordan.
1980-1981 Part-time Lecturer, Department of Biology, American University of Beirut, Beirut, Lebanon.
1978-1980 Supervising Laboratory Courses in Biochemistry to final year honors students at the University of Cambridge, Cambridge, United Kingdom.
1976-1977 Lecturer, Department of Biological Sciences, Yarmouk University, Irbid, Jordan.
1971-1974 Instructor, Department of Biological Sciences, University of Jordan, Amman, Jordan.

Professional Affiliations

- Cambridge Philosophical Society, United Kingdom.
- The Biochemical Society, United Kingdom.
- American Society of Plant Biologists, USA.
- Fellow of the Institute of Biology, London, United Kingdom.
- Fellow of the Third World Academy of Sciences (TWAS), Trieste, Italy.
- Founding Fellow and Secretary General, Arab Academy of Sciences, Beirut, Lebanon.
- Honorary Research Fellow, Institute of Biomedical and Life Sciences, University of Glasgow, United Kingdom.
- The Darwin Society, Darwin College, Cambridge, United Kingdom.
- The Lebanese Association for the Advancement of Sciences, Lebanon